SUBGENUS Pleurothallis SECTION Macrophyllae-Fasciclatae Lindl. 1859

Characterized by the single flowered inflorescence with the peduncle hidden within a spathe, the usually short pedicels, the entire, fringed to pubescent petals that are not sigmoid in outline and a thick, obtuse, oblong to transverse lip and a sorter, broader column with an apical anther and stigma that is almost always bilobed. This subsection and amphygiae are not distinctly separated in various borderline species and for this reason they later were separated by Luer into the genus Acronia [2005] with three sections, Acronia, Amphygia and Macrophyllae-fasciculatae. TYPE Species for the SECTION

!Pleurothallis calceolaris Rchb.f. 1876

  1. Pleurothallis acestrophylla Luer 1975 Ecuador and Peru

  2. Pleurothallis acutilabia Luer 1999 Colombia

  3. Pleurothallis adeleae Luer 1981 Ecuador and Ecuador

  4. Pleurothallis adelphe Luer & Hirtz 1996 Ecuador

  5. Pleurothallis adonis Luer 1976 Colombia and Ecuador

  6. Pleurothallis adventurae Karremans & Bogarín 2011 Costa Rica

  7. Pleurothallis aguirrei Luer & R.Escobar 2003 Colombia

  8. Pleurothallis alopex Luer 1976 Bolivia and Paraguay

  9. Pleurothallis altimonile Luer & R.Escobar 1981 Colombia

  10. Pleurothallis ambyx Luer & R.Escobar 1981 Colombia and Ecuador

  11. Pleurothallis andreettae (Luer) J.M.H.Shaw 2014 Ecuador

  12. Pleurothallis angusta Ames & C.Schweinf. 1925 Costa Rica and Panama

  13. Pleurothallis angustissima (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

  14. Pleurothallis anthurioides A.Doucette 2016 Panama

  15. Pleurothallis apopsis Luer 1979 Ecuador

  16. Pleurothallis applanata Luer & Dalström 1996 Ecuador

  17. Pleurothallis archicolonae Luer 1976 Panama

  18. Pleurothallis arctata Luer 1997 Peru

  19. Pleurothallis ascera Luer & R. Escobar 1996 Costa Rica, Panama and Colombia

  20. Pleurothallis asplundii Luer 1996 Ecuador

  21. Pleurothallis aurita Schweinf 1940 Costa Rica

  22. Pleurothallis baezensis (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

  23. Pleurothallis batrachus Luer & Hirtz 1996 Ecuador

  24. Pleurothallis baudoensis Luer & R.Escobar 1998 Colombia

  25. Pleurothallis belocardia Schltr. 1924 Colombia

  26. Pleurothallis bitumida Luer 1996 Costa Rica

  27. Pleurothallis bivalvis Lindl. 1846 Ecuador, Colombia and Venezuela

  28. Pleurothallis blepharopetala Schltr.1915 Ecuador

  29. Pleurothallis bogarinii Pupulin & J.D. Zúñiga 2007 Costa Rica

  30. Pleurothallis bothros Luer 1996 Costa Rica

  31. Pleurothallis bovilingua Luer & R.Escobar 1998 Colombia

  32. Pleurothallis braidiana [Luer] in ed. without location

  33. Pleurothallis bucculenta Luer 1976 Ecuador

  34. Pleurothallis bucranon Luer & Hirtz 1988 Ecuador

  35. Pleurothallis bulbosa [Luer & Escobar] Pfahl 2012 Colombia

  36. Pleurothallis calamifolia Luer & R.Escobar 1996 Colombia

  37. !Pleurothallis calceolaris Rchb.f. 1876 Colombia

  38. Pleurothallis callifera C.Schweinf. 1951 Venezuela

  39. Pleurothallis calogramma Luer 2003 Colombia

  40. Pleurothallis callosa Diaz & Pupulin 2021 Costa Rica

  41. Pleurothallis calolalax Luer & R.Escobar 1998 Colombia

  42. Pleurothallis canaligera Rchb.f 1877 Colombia, Ecuador and Peru

  43. Pleurothallis canidentis Luer & R. Escobar 1998 Costa Rica, Panama and Colombia

  44. Pleurothallis cardiochila L.O. Williams 1942 Costa Rica and Panama

  45. Pleurothallis cardiocrepis Rchb.f.1888 Peru?

  46. Pleurothallis cardiophylax Rchb.f. 1888 Colombia, Ecuador and Peru

  47. Pleurothallis cardiostola Rchb. f. 1854 Colombia, Venezuela, Peru, Bolivia and Ecuador

  48. Pleurothallis cardiothallis Rchb.f 1857 Mexico, Belize, Costa Rica, Nicaragua, Colombia and Ecuador

  49. Pleurothallis carduela (Luer) J.M.H.Shaw 2014 Ecuador

  50. Pleurothallis carnosa Braid 1924 Colombia

  51. Pleurothallis carrenoi Carnevali & I.Ramírez 1987 Venezuela

  52. Pleurothallis cassidata Luer & R.Escobar 1998 Colombia

  53. Pleurothallis castanea Mark Wilson, G.Merino & J.D.Werner 2016 Ecuador

  54. Pleurothallis cauda-hirundinis Luer & J.Portilla 2002 Ecuador

  55. Pleurothallis cedrinorum Luer & Dalström 1996 Ecuador

  56. Pleurothallis chama Luer 1977 Ecuador

  57. Pleurothallis chaparensis (Luer & R.Vásquez) J.M.H.Shaw 2014 Bolivia

  58. Pleurothallis chavezii Luer 1996 Costa Rica

  59. Pleurothallis chicalensis M.M.Jiménez & Baquero 2018 Ecuador

  60. Pleurothallis cobriformis L.O.Williams 1940 Panama

  61. Pleurothallis complanata Luer & Hirtz 1996 Ecuador

  62. Pleurothallis compressa Luer 1996 Costa Rica

  63. Pleurothallis conformalis Luer & Dalström 2003 Ecuador

  64. Pleurothallis conicostigma Luer 1975 Ecuador

  65. Pleurothallis constricta Luer & R.Escobar R. 1998 Colombia

  66. Pleurothallis convexa Luer & Hirtz 1996 Ecuador

  67. Pleurothallis cordata (Ruiz & Pav.) Lindl. 1830 Colombia, Venezuela, Ecuador, Peru and Bolivia

  68. Pleurothallis cordifolia Rchb.f. & Wagener 1854 Costa Rica, Panama, Venzuela, Colombia and Ecuador

  69. Pleurothallis coriacardia Rchb. f. 1854 Ecuador and Peru

  70. Pleurothallis correllii Luer 1976 Guatemala, El Salvador and Mexico

  71. Pleurothallis corysta Luer 1978 Ecuador

  72. Pleurothallis crateriformis C.Schweinf. 1951 Ecuador and Peru

  73. Pleurothallis crossota Luer & Dalström 1996 Ecuador

  74. Pleurothallis crucifera Luer & Hirtz 1996 Ecuador

  75. Pleurothallis culpameae (Luer) J.M.H.Shaw 2014 Ecuador

  76. Pleurothallis cuzcoensis [Luer] ined. Peru

  77. Pleurothallis cutucuensis Luer 1977 Ecuador

  78. Pleurothallis cyanea Luer & R. Escobar 1984 Colombia

  79. Pleurothallis deflexa Luer 1975 Ecuador

  80. Pleurothallis dejavu Luer & Hirtz 1988Ecuador

  81. Pleurothallis depressa Luer & R. Escobar 1996 Colombia

  82. Pleurothallis dewildei Luer & R.Escobar 1998 Colombia

  83. Pleurothallis diabolica Luer & R. Escobar 1981 Colombia Ecuador

  84. Pleurothallis diazii (Luer & Endara) J.M.H.Shaw 2014 Peru

  85. Pleurothallis dibolia Luer 1975 Ecuador

  86. Pleurothallis dilemma Luer 2001 Ecuador

  87. Pleurothallis discoidea Lindl. 1835 Trinidad, Venezuela, Surinam, French Guiana, Ecuador and Peru

  88. Pleurothallis dorotheae Luer 1980 Costa Rica

  89. Pleurothallis dracuncula (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

  90. Pleurothallis drewii Luer 2000

    Ecuador

  91. Pleurothallis dubbeldamii Luer 2003 No Collection Data

  92. Pleurothallis ensata Luer 1975 Ecuador

  93. Pleurothallis epiglottis Luerb 1980 Ecuador

  94. Pleurothallis erythrium Luer 1978 Ecuador

    Pleurothallis escobarii (Luer) J.M.H.Shaw 2014 Colombia

  95. Pleurothallis excavata Schltr. 1923 Costa Rica and Panama

  96. Pleurothallis fantastica Ames 1936 Costa Rica

  97. Pleurothallis folsomii (Luer & Endara) J.M.H.Shaw 2014 Panama

  98. Pleurothallis fonnegrae (Luer & Endara) J.M.H.Shaw 2014 Colombia

    ~

  99. Pleurothallis fossulata Luer & R.Escobar 1998 Colombia and Ecuador

  100. Pleurothallis galerita Luer 1981 Ecuador

  101. Pleurothallis ganymedes Luer & R.Escobar 1981 Colombia

  102. Pleurothallis garciae Luer 1999 Venezuela

  103. Pleurothallis gargantua Luer 1996 Ecuador

  104. Pleurothallis geographica Luer 1977 Ecuador

  105. Pleurothallis giraffa Luer 1979 Ecuador

  106. Pleurothallis giraldoi Luer 1999 Colombia

  107. Pleurothallis glabra Luer & R.Escobar 1996 Colombia

  108. Pleurothallis globosa Luer & R.Escobar 1996 Colombia

  109. Pleurothallis glochis Luer & R.Escobar 1981 Colombia and Ecuador

  110. Pleurothallis gomezii Luer & R.Escobar 1996 Colombia

  111. Pleurothallis gonzaleziorum Pupulin, Diaz & Pridgeon 2021 Costa Rica

  112. Pleurothallis grandiflora Lindl. 1835 Venezuela, Colombia, Ecuador, Peru and Bolivia

  113. Pleurothallis grandilingua Pupuplin, M Diaz and Pridgeon 2021 Costa Rica

  114. Pleurothallis hemileuca Luer 1982 Panama

  115. Pleurothallis hoeijeri Luer & Hirtz 1988 Ecuador

  116. Pleurothallis homalantha Schltr.1923 El Salvador, Honduras, Nicaragua, Costa Rica, Panama and Brazil

  117. Pleurothallis imitor Luer 1996 Costa Rica

  118. Pleurothallis imperialis Luer 1975 Ecuador and Colombia

  119. Pleurothallis index Luer 1980 Ecuador and Colombia

  120. Pleurothallis isthmica Luer 1977 Honduras, Costa Rica and Panama

  121. Pleurothallis jaramilloi Luer 2001 Colombia

  122. Pleurothallis jupiter Luer 1975 Ecuador

  123. Pleurothallis killipii Garay 1956 Colombia

  124. Pleurothallis lacera Luer 1975 Colombia and Ecuador

  125. Pleurothallis languida Luer & R.Escobar 1998 Colombia

  126. Pleurothallis latipetala Garay 1952 Colombia

  127. Pleurothallis lemniscifolia Luer 1975 Ecuador

  128. Pleurothallis leucantha Schltr. 1912 Guatemala

  129. Pleurothallis lilijae Foldats 1968 Venezuela, Colombia and Ecuador

  130. Pleurothallis linguifera Lindl. 1859 Colombia, Ecuador, Peru, Bolivia and Venezuela

  131. Pleurothallis longipetala Bogarin & Belfort 2021 Costa Rica

  132. Pleurothallis loreae Carnevali & I.Ramírez 2003 Venezuela

  133. Pleurothallis luna-crescens Pupulin, J.Aguilar & Mel.Fernández 2017 Costa Rica

  134. Pleurothallis lunaris Luer & R.Escobar 1981 Colombia

  135. Pleurothallis lynniana [Luer] Comb. novo. Pfahl 2012 Ecuador

  136. Pleurothallis macra Lindl. 1859 Ecuador and Peru

  137. Pleurothallis macrocardia Rchb.f. 1855 Colombia, Ecuador and Bolivia

  138. Pleurothallis maduroi Luer 1997 Panama

  139. Pleurothallis magnipetala C.Schweinf. 1951 Peru

  140. Pleurothallis marioi Zambrano & Solano 2017 Ecuador

  141. Pleurothallis marthae Luer & R. Escobar 1996 Colombia

  142. Pleurothallis mastodon Luer & Hirtz 1996 Ecuador

  143. Pleurothallis matudana C. Schweinf. 1938 Mexico, Guatemala, Honduras, Nicaragua, Ecuador and Bolivia

  144. Pleurothallis megaglossa Luer & Dalström 1996 Ecuador

  145. Pleurothallis megalorhina Luer & R.Escobar 1996 Colombia

  146. Pleurothallis mesopotamica Bogarin & Belfort 2021 Costa Rica

  147. Pleurothallis microcardia Rchb.f. 1855 Colombia, Venezuela, Ecuador, Peru and Bolivia

  148. Pleurothallis millei Schltr. 1915 Ecuador

  149. Pleurothallis minutilabia Mark Wilson, Tobar & Á.J.Pérez 2018 Ecuador

  150. Pleurothallis monocardia Rchb. f. 1855 Venezuela, Colombia and Ecuador

    ~

  151. Pleurothallis muriculata Luer & Hirtz 1996 Ecuador

  152. Pleurothallis navisepala Pupulin, J.Aguilar & M.Díaz 2017 Costa Rica

  153. Pleurothallis neorinkei A.Doucette 2016 Colombia

  154. Pleurothallis neossa (Luer & Hirtz) J.M.H.Shaw 2014 Ecuador

  155. Pleurothallis nephroglossa Schltr. 1921 Ecuador

  156. Pleurothallis nipterophylla C.A.Luer 1976 Ecuador

  157. Pleurothallis nitida Luer 1977 Panama

  158. Pleurothallis nossax Luer & R. Escobar 1981 Colombia

  159. Pleurothallis octavioi Luer & R.Escobar R. 1981 Colombia

  160. Pleurothallis omoglossa Luer 1976 Ecuador

  161. Pleurothallis oncoglossa Luer 1996 Costa Rica

  162. Pleurothallis ortegae Luer & Hirtz 1996 Ecuador

  163. Pleurothallis oscarii Archila & Chiron 2011 Guatemala

  164. Pleurothallis oxapampae Luer 1982 Peru

  165. Pleurothallis pallida Luer 1977 Panama

  166. Pleurothallis palliolata Ames 1922 Costa Rica and Panama

  167. Pleurothallis pansamalae Schltr. 1912 Mexico, Guatemalla, Belize, El Salvador, Honduras and Nicaragua

  168. Pleurothallis paquishae Luer 1996 Ecuador and Peru

  169. Pleurothallis peculiaris Luer 1976 Panama

  170. Pleurothallis pelicophora Luer 2001 Ecuador

  171. Pleurothallis penelops Luer 1978 Ecuador

  172. Pleurothallis perijaensis Dunst. 1978 Venezuela, Colombia and Ecuador

  173. Pleurothallis peroniocephala Luer 1975 Ecuador

  174. Pleurothallis perryi Luer 1979 Ecuador

  175. Pleurothallis persimilis Luer & Hirtz 2003 Ecuador

  176. Pleurothallis phyllocardia Rchb.f. 1866 Costa Rica and Panama

  177. Pleurothallis phyllocardioides Schltr. 1923 Guatemala, Nicaragua, Costa Rica, Panama, Colombia, Ecuador, Peru, Bolivia and Venezuela

  178. Pleurothallis phymatodea Luer 1976 Ecuador

  179. Pleurothallis pileata Luer & R.Escobar 1982 Colombia

  180. Pleurothallis platypetala Luer & R.Escobar 1996 Colombia

  181. Pleurothallis platysepala Schltr. 1920 Colombia

  182. Pleurothallis portillae Luer 1979 Colombia and Ecuador

  183. Pleurothallis prolaticollaris Luer 1977 Ecuador

  184. Pleurothallis pseudopogon Luer & Hirtz 1998 Ecuador

  185. Pleurothallis pudica Pupulin, J.Aguilar & M.Díaz 2017 Costa Rica

  186. Pleurothallis pyelophera [Luer] comb. novo. Pfahl 2013 Ecuador

  187. Pleurothallis quitu-cara Carrera & Baquero 2018 Ecuador

  188. Pleurothallis radula Luer 1979 Costa Rica

  189. Pleurothallis ramosii Luer 1998 Colombia

  190. Pleurothallis rectipetala Ames & Schweinf. 1925 Panama

  191. Pleurothallis recurvata Luer & Hirtz 1996 Ecuador

  192. Pleurothallis reptans Luer 1981 Ecuador

  193. Pleurothallis rhinocera (Luer & Sijm) J.M.H.Shaw 2013 Ecuador

  194. Pleurothallis rhodoglossa Schltr.1922 Costa Rica, Panama, Colombia and Ecuador

  195. Pleurothallis ripleyi Luer 1979 Ecuador

  196. Pleurothallis ruberrima Lindley 1846 Colombia, Ecuador, Peru, Bolivia and Venezuela

  197. Pleurothallis rubrifolia Mark Wilson, Tobar & Salas Guerr. 2016 Ecuador, Peru

  198. Pleurothallis rubroinversa Luer 1975 Ecuador

  199. Pleurothallis rugosa Luer & R.Escobar 1982 Colombia

  200. Pleurothallis saccatilabia C.Schweinf. 1940 Mexico and Guatemala

  201. Pleurothallis sagittilabia Luer 1998 Colombia

  202. Pleurothallis sanchoi Ames 1923 Guatemala, Costa Rica and Panama

  203. Pleurothallis sannio Luer & R.Escobar 1988 Colombia

  204. Pleurothallis sarcochila Garay 1953 Ecuador

  205. Pleurothallis scabrilinguis Lindl. 1836 Colombia Ecuador and Peru

  206. Pleurothallis scaphipetala Luer 1996 Costa Rica

  207. Pleurothallis scotinantha Pupulin, M.Díaz & J.Aguilar 2017 Costa Rica

  208. Pleurothallis scurrula Luer 1976 Peru

  209. Pleurothallis semiscabra Lindl. 1846 Venezuela

  210. Pleurothallis sergioi Luer & R.Escobar 2003 Colombia

  211. Pleurothallis serricardia Schltr. 1924 Colombia

  212. Pleurothallis sigynes Luer 1976 Colombia & Ecuador

  213. Pleurothallis silvae-pacis Karremans 2011 Costa Rica

  214. Pleurothallis siphoglossa Luer & R. Escobar 1996 Colombia

  215. Pleurothallis sotarae Schltr. 1920 Colombia

  216. Pleurothallis sphaerantha Luer 1975 Ecuador

  217. Pleurothallis stelidilabia Luer 1979 Ecuador

  218. Pleurothallis stellata I.Ramírez & Carnevali 2003 Venezuela

  219. Pleurothallis stenosepala Rolfe 1892 Venezuela, Colombia, Ecuador, Peru and Bolivia

  220. Pleurothallis stenota Luer 1977 Colombia and Ecuador

  221. Pleurothallis subtilis C.Schweinf. 1951 Venezuela and Colombia

  222. Pleurothallis x subversa Pupulin & Bogarin 2021 Costa Rica

  223. Pleurothallis tamaensis Foldats 1968 northwestern Venezuela

  224. Pleurothallis tapatiensis Pupulin, M Diaz & Pridgeon 2021 Costa Rica

  225. Pleurothallis teaguei Luer 1996 Ecuador

  226. Pleurothallis titan Luer 1977 Panama and Colombia

  227. Pleurothallis tobarii [Luer & Hirtz] comb. novo. Pfahl 2012 Ecuador

  228. Pleurothallis tonduzii Schltr. 1918 Costa Rica, Panama and Peru?

  229. Pleurothallis tragulosa Luer & Hirtz 1996 Ecuador

  230. Pleurothallis transversilabia Carnevali & I.Ramírez 2000 Venezuela

    Pleurothallis triangulilabia C.Schweinf. 1937 Costa Rica

  231. Pleurothallis tridentata Klotzsch 1840 Venezuela, Colombia, Ecuador, Peru? and Bolivia?

  232. Pleurothallis troglodytes Luer 1982 Ecuador

  233. Pleurothallis tyria Luer & Hirtz 1996 Ecuador

  234. Pleurothallis undulata Poepp. & Endl. 1836 Peru

  235. Pleurothallis upanoensis Luer & Hirtz 2003 Ecuador

  236. Pleurothallis valladolidensis Luer 1983 Ecuador

  237. Pleurothallis variabilis Luer 1979 Colombia and Ecuador

  238. Pleurothallis vinealis Luer & R.Escobar 1986 Colombia and Ecuador

  239. Pleurothallis volans Luer & Hirtz 1996 Ecuador

  240. Pleurothallis zarumae Luer & Hirtz 1996 Ecuador

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